2018 Jul. – 2019 Jan., Postdoctoral researcher in Department of Paleobiology, National Museum of Natural History, Smithsonian Institution. Supervisor: Douglas H. Erwin (Senior Scientist and Curator).

2018 Jul. – 2018 Aug., Trainee in Analytical Paleobiology Workshop, University of Florida. Organizing Chair: Michal Kowalewski (Professor).

2016 Sep. – 2018 Mar., Predoctoral researcher in Department of Paleobiology, National Museum of Natural History, Smithsonian Institution. Supervisor: Douglas H. Erwin (Senior Scientist and Curator).

2013 Sep. – 2018 Jun., Ph.D. Degree of Paleontology and Stratigraphy, University of Chinese Academy of Sciences. Studied in Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences. Supervisor: Maoyan Zhu (Professor), Fangchen Zhao (Professor).

2009 Sep. – 2013 Jul., Bachelor Degree of Geology (with focus on Paleontology), Nanjing University.

　　My current research focuses on the origin and early evolution of arthropods, segmented and legged invertebrates and the most diverse animal group on the Earth ever since the Cambrian Explosion. I have been investigating the vast collections of soft-bodied arthropods from the exceptionally preserved fossil deposits in the Cambrian (e.g. Chengjiang, Burgess Shale, etc., 520–505 million years ago) for years. These fossils document invaluable morphologies of early arthropods and their close relatives. Therefore, I am devoted to rereading the early evolutionary history of arthropods by integrating perspectives from paleobiology, phylogeny and development.

1. Zeng H, Zhao FC, Zhu MY. 2022. Innovatiocaris, a complete radiodont from the early Cambrian Chengjiang Lagerst？tte and its implications for the phylogeny of Radiodonta. Journal of the Geological Society, 180(1): jgs2021-164.

2. 曾晗. 2022. 节肢动物起源难题等待解答. 前沿科学, 16(63): 16–21.

3. Zeng H, Zhao FC, Yin ZJ, Zhu MY. 2021. A new early Cambrian bivalved euarthropod from Yunnan, China and general interspecific morphological and size variations in Cambrian hymenocarines. Palaeoworld, 30(3), 387–397.

4. Zeng H, Zhao FC, Niu KC, Zhu MY, Huang DY. 2020. An early Cambrian euarthropod with radiodont-like raptorial appendages. Nature, 588(7836): 101–105.

5. Zeng H, Zhao FC, Yin ZJ, Zhu MY. 2018. Morphology of diverse radiodontan head sclerites from the early Cambrian Chengjiang Lagerst？tte, south-west China. Journal of Systematic Palaeontology, 16(1): 1–37.

6. Zeng H, Zhao FC, Yin ZJ, Zhu MY. 2018. A new radiodontan oral cone with a unique combination of anatomical features from the early Cambrian Guanshan Lagerst？tte, eastern Yunnan, South China. Journal of Paleontology, 92(1): 40–48. (封面论文)

7. Zeng H, Zhao FC, Yin ZJ, Zhu MY. 2017. Appendages of an early Cambrian metadoxidid trilobite from Yunnan, SW China support mandibulate affinities of trilobites and artiopods. Geological Magazine, 154(6): 1306–1328.

8. Zeng H, Zhao FC, Yin ZJ, Li GX, Zhu M. 2014. A Chengjiang-type fossil assemblage from the Hongjingshao Formation (Cambrian Stage 3) at Chenggong, Kunming, Yunnan. Chinese Science Bulletin, 59(25): 3169–3175. (封面论文)

9. Tian QY, Zhao FC, Zeng H, Zhu MY, Jiang BY. 2022. Ultrastructure reveals ancestral vertebrate pharyngeal skeleton in yunnanozoans. Science, 377(6602): 218–222.

10. Sun ZX, Zhao FC, Zeng H. 2022. Unique Cambrian trilobite Eodontopleura: an example of strong convergence in trilobite evolution, Historical Biology, doi: 10.1080/08912963.2022.2157273.

11. Sun ZX, Zhao FC, Zeng H, Luo C, Van Iten H, Zhu MY. 2022. The middle Cambrian Linyi Lagerst？tte from the North China Craton: a new window on Cambrian evolutionary fauna. National Science Review, 9: nwac069.

12. Wang DZ, Zhao FC, Zeng H, Li GX, Zhu MY. 2022. A new rhynchonelliform brachiopod Longtancunella with soft-part preservation from the Hongjingshao Formation (Cambrian Stage 3) in Yunnan, South China. Palaeoworld, 31(4), 570–581.

13. 孙智新, 曾晗, 赵方臣. 2021. 山东寒武系苗岭统三叶虫消化系统的化石证据. 古生物学报, 60(1): 2020024.

14. 刘瑶, 孙智新, 曾晗, 赵方臣. 2021. 山东潍坊寒武系苗岭统乌溜阶馒头组上部的腕足动物. 古生物学报, 60(1): 2020051.

15. Sun ZX, Zeng H, Zhao FC. 2020. A new middle Cambrian radiodont from North China: Implications for morphological disparity and spatial distribution of hurdiids. Palaeogeography, Palaeoclimatology, Palaeoecology 558: 109947.

16. Sun ZX, Zeng H, Zhao FC. 2020. Occurrence of the hurdiid radiodont Cambroraster in the middle Cambrian (Wuliuan) Mantou Formation of North China. Journal of Paleontology 94(5): 881–886.

17. Sun ZX, Zeng H, Zhao FC. 2020. First occurrence of the Cambrian arthropod Sidneyia Walcott, 1911 outside of Laurentia. Geological Magazine 157(3): 405–410.

18. Sun ZX, Zeng H, Zhao FC. 2020. A new middle Cambrian trilobite with a specialized cephalon from Shandong Province, North China. Acta Palaeontologica Polonica 65(4): 709–718.

19. Aria C, Zhao FC, Zeng H, Guo J, Zhu MY. 2020. Fossils from South China redefine the ancestral euarthropod body plan. BMC Evolutionary Biology 20(4): 1–17.

20. Luo C, Zhao FC, Zeng H, 2020. The first report of a vauxiid sponge from the Cambrian Chengjiang Biota. Journal of Paleontology, 94(1): 28–33.

21. Sun HJ, Yin ZJ, Li GX, Zhao FC, Zeng H, Zhu MY. 2020. Periodic shell decollation as an ecology-driven strategy in the early Cambrian Cupitheca. Palaeontology, 63(3): 431–442.

22. Sun HJ, Zhao FC, Steiner M, Li GX, Na L, Pan B, Yin ZJ, Zeng H, Van Iten H, Zhu MY. 2020. Skeletal faunas of the lower Cambrian Yu'anshan Formation, eastern Yunnan, China: Metazoan diversity and community structure during the Cambrian Age 3. Palaeogeography, Palaeoclimatology, Palaeoecology, 542: 109580.

23. 朱茂炎, 赵方臣, 殷宗军, 曾晗, 李国祥. 2019. 中国的寒武纪大爆发研究: 进展与展望. 中国科学: 地球科学, 49(10): 1455–1490.

24. Sun HJ, Smith M, Zeng H, Zhao FC, Li GX, Zhu MY. 2018. Hyoliths with pedicles illuminate the origin of the brachiopod body plan. Proceedings of the Royal Society B: Biological Sciences, 285(1887): 20181780.

25. Sun HJ, Zhao FC, Wen RQ, Zeng H, Peng J. 2018. Feeding strategy and locomotion of Cambrian hyolithides. Palaeoworld, 27(3): 334–342.

26. Yin ZJ, Zhao DD, Pan B, Zhao FC, Zeng H, Li GX, Bottjer DJ, Zhu MY. 2018. Early Cambrian animal diapause embryos revealed by X-ray tomography. Geology, 46(5): 387–390.

27. Smith EF, Nelson LL, Tweedt SM, Zeng H, Workman JB. 2017. A cosmopolitan late Ediacaran biotic assemblage: new fossils from Nevada and Namibia support a global biostratigraphic link. Proceedings of the Royal Society B: Biological Sciences, 284(1858): 20170934.

28. Zhao FC, Smith M, Yin ZJ, Zeng H, Li GX, Zhu MY. 2017. Orthrozanclus elongata n. sp. and the significance of sclerite-covered taxa for early trochozoan evolution. Scientific Reports, 7(1): 16232.

29. Zhao FC, Smith M, Yin ZJ, Zeng H, Hu SX, Li G, Zhu MY. 2015. First report of Wiwaxia from the Cambrian Chengjiang Lagerst？tte. Geological Magazine, 152(2): 378–382.

30. Zhao FC, Hu SX, Zeng H, Zhu MY. 2014. A new helmetiid arthropod from the early Cambrian Chengjiang Lagerst？tte, southwest china. Journal of Paleontology, 88(2): 367–370.

科普论文及奖项

1. 曾晗. 2021. 揭开“节肢动物起源”之谜的关键过渡型化石——麒麟虾. 大自然, 202102(218): 26–29. (科普文章, 杂志封面)

2. 曾晗. 2021. 解密节肢动物起源——麒麟虾. 知识就是力量, 202109(586): 6–7. (科普文章)

3. 盛捷, 曾晗, 蔡秀伟, 陈聪. 2021. 五亿年前奇特的“麒麟虾”——揭秘节肢动物起源的过渡型化石. (科普视频)

4. 2020年全国优秀科普微视频作品。

5. 2020年中国科学院优秀科普微视频。

6. 2020年江苏省第十二届优秀科普作品影视动画类一等奖。